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Kratos is revealed to have been gathering materials to have Lloyd use the Eternal Sword. They decide to confront and defeat Mithos before seeking the Eternal Sword.

Their actions instead destabilize the Great Seed, the supplier of mana to both worlds, causing it to grow and engulf Sylvarant at an exponential rate. After the party re-stabilizes the seed, they learn from Yuan Ka-Fai about the origins of Sylvarant and Tethe'alla and how they used to be one world but was split by Mithos with the Eternal Sword the party realizes they can save the world if they wield the Eternal Sword, merge the worlds, and germinate the Great Seed. The party learns that awakening the Summon Spirits in both worlds will sever the mana tie between them. Believing that to be the answer, they successfully sever all mana ties between the two worlds. The party leaves for Tethe'alla to search for a way to save both worlds and are joined by Zelos Wilder, Presea Combatir, and Regal Bryant. She tells them about how the two worlds vie for each other's mana through the journey of regeneration. Hoping to find answers from the angel Remiel, they party meets him at the final seal and are betrayed by Remiel and Kratos both are revealed to be part of Cruxis, an evil organization led by Mithos Yggdrasill.

On their journey, they meet Sheena Fujibayashi who comes from Tethe'alla, a world parallel to Sylvarant. The purpose of the journey is to replenish Sylvarant with mana, a necessary energy to their survival. The path of the journey consists of five temples which must be unsealed by Colette. Lloyd Irving and his friend, Genis Sage accompany the chosen, Colette Brunel and her guardians, Raine Sage and Kratos Aurion on her journey of world regeneration.

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